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Many generations after Caesar's death, apes have become the dominant species on Earth, establishing numerous clans, while the surviving humans have become feral. Noa, a young chimpanzee hunter from a falconry-practicing clan, prepares for a coming-of-age ceremony by collecting wild eagle eggs with his friends Anaya and Soona. However, a human scavenger follows Noa home and inadvertently cracks his egg during a scuffle before fleeing. While searching for a replacement egg, Noa encounters a group of ape raiders using electric weapons. While Noa hides from them, the apes follow his horse back to his clan. Noa hurries home to find his village burning; the gorilla general Sylva kills Noa's father, Koro, before dropping Noa from a high platform. Noa awakens to find that his clan has been abducted. He buries Koro and sets out to rescue his clan. Noa is joined by Raka, an orangutan who tells Noa about Caesar's teachings. The apes notice they are being followed by the human scavenger; Raka offers her food and a blanket, naming her Nova. When the trio encounters a group of feral humans with a herd of zebras, Sylva's raiders suddenly attack them. Noa and Raka rescue Nova, who, to their surprise, is intelligent and can speak. She reveals that her name is Mae and that the raiders took Noa's clan to a beachfront settlement outside an old human vault. As they cross a bridge on their way to the settlement, they are ambushed by Sylva. In the ensuing fight, Raka saves Mae from drowning but is swept away by the rapids. Noa and Mae are captured and taken to the apes' settlement. Noa reunites with his clan and is introduced to the apes' king, Proximus Caesar. Proximus has enslaved other clans, forcing them to work on opening the vault so he can access the human technology locked inside. Proximus invites Noa to dinner with Mae and Trevathan, another speaking human prisoner who teaches Proximus about the old human world. Proximus believes that Noa's intuitiveness could help open the vault and warns him that Mae only has her own agenda. Noa confronts Mae, demanding the truth in exchange for his help. Mae discloses knowledge of a hidden entrance to the vault and seeks a book capable of restoring speech to humanity. Noa agrees to help her enter and destroy the vault. Noa, Mae, Soona, and Anaya secretly plant explosives around the sea walls that protect the settlement. Trevathan catches them and intends to warn Proximus, but Mae strangles him to death. The group enters the vault, finding a stockpile of weapons and Mae's "book", which is actually a satellite decryption key. The apes discover old children's picture books depicting humans as the once-dominant species and apes in cages at a zoo. As the group makes their way out of the bunker, they are confronted by Proximus and his followers. Lightning, one of Proximus' lieutenants, threatens to kill Soona, but Mae kills him with a gun and triggers the explosives, flooding the bunker with the apes inside. Mae flees the settlement while the apes climb to higher ground through the bunker. Noa is pursued by Sylva, whom he traps and leaves to drown. Noa then escapes the bunker but is attacked by Proximus. Noa leads his clan in summoning their eagles to attack Proximus and send him falling off a cliff into the sea. As Noa's clan returns to rebuild their home, Mae arrives to bid farewell to Noa, secretly holding a gun behind her back. She explains that she destroyed the bunker to prevent the apes from obtaining its weapons. Given their disparities, Noa and Mae both question whether apes and humans can co-exist peacefully. As Noa takes Soona to look through a telescope he found on his journey, Mae travels to an intelligent human settlement at a satellite base. Mae delivers the decryption key, allowing them to reactivate the satellites and successfully contact other intelligent humans worldwide.